Publications — The Levi Lab

Comparative foraging ecology of lynx and coyotes in the Gulf of Alaska national parks

Dymit, E., Griffin, K., Morton, E., Hower, J., Levi, T.

Apparent phenotypic mismatch limits the recovery of a reintroduced carnivore

Barry, B., Smith, G., Eriksson, C., Moriarty, K., Hartman, J., et al.

Spatial Memory and Resource Selection Patterns in Translocated African Elephants

Ngosi, M., Ruprecht, J.S., Lesmeister, D.B., Levi, T.

Assessing unmarked models for estimating densities of small mammals using camera traps

Tosa, M., Weldy, M., Epps, C., Lesmeister, D., Levi, T.

Ecological Applications

Listening for extinction: Passive acoustic monitoring highlights critical risks and priorities for northern spotted owls

Lesmeister, D.B., Chestnut, T., Ellis, T., et al.

Are occupancy models always appropriate? Aligning state variables, model assumptions, and study objectives

Twining, J., Barnas, A., Burton, C., et al.

Leveraging acoustic bycatch to identify and map the distribution of a temperate forest sentinel

Kohlberg, A., Levi, T., Hutchinson, R.A., Kim, J.B., Lesmeister, D.B.

Comparing accuracy and biases of DNA metabarcoding, hybridization capture, and metagenomic sequencing for quantifying herbivore diets

Eriksson, C.E., Shipley, L., Clark, D.A., Levi, T.

Metabarcoding versus parasite-specific real-time PCRs in the detection of nematodes of veterinary importance

Robleto-Quesada, J., Benavides, G., Solano-Barquero, A., et al.

The effects of predator recovery and climate change on the long-term demography of a large herbivore

Loonam, K., Brown, C., Clark, D., Rowland, M.M., Wisdom, M., Levi, T.

Leveraging next-generation tools for genetic assessment and demographic monitoring in threatened and elusive Humboldt martens

Hallerud, M.A., Moriarty, K., Pilgrim, K.L., Quinn, C.B., Eriksson, C.E., Schwartz, M.K., Levi, T.

Evolutionary Applications 19(7):e70277

Individual identification of prey in carnivore scats

Eriksson, C.E., Bianco, M., Ruprecht, J., Clark, D.A., Levi, T.

Molecular Ecology Resources 26(5):e70164

Drought and growing season phenology over 35 years modulates species interactions among domestic and wild herbivores

Ruprecht, J., Forrester, T.D., Brown, C.L., Mengak, L.F., Wisdom, M.J., Clark, D.A., Levi, T.

Journal of Animal Ecology

Diets, dominance hierarchies, and kleptoparasitism drive asymmetrical interactions between wolves and cougars

Binder, W., Ruprecht, J.S., Rabe, J., Metz, M.C., Hutchinson, R., Stahler, D.R., Levi, T.

Proceedings of the National Academy of Sciences

Symbolic regression for empirically realistic population dynamic time series

Jarman, C.N., Levi, T., Novak, M.

bioRxiv (preprint)

PNW-Cnet: An evolving convolutional neural network to support broad-scale passive acoustic monitoring

Lesmeister, D.B., Ruff, Z.J., Duarte, A., Kohlberg, A.B., Levi, T., Sullivan, C.M.

Ecological Informatics

Large-scale metagenomic surveillance study expands the known diversity of RNA viruses in mosquito populations from the Amazon Basin

Fuques, E., Massey, A.L., Qureshi, F., Campos-Silva, J.V., da Silva, D.J.F., Levi, T.

PeerJ

Vertebrate community composition and activity at giant otter latrines in the northern Pantanal

Olden, E.P., Eriksson, C.E., Kantek, D.L.Z., Miyazaki, S.S., dos Santos-Filho, M., Muniz, C.C., Campos, D.V.S., Peres, C.A., Levi, T.

Ecosphere 17(4):e70619

Prey Partitioning in a Diverse Carnivore Community: Implications for Reintroduced Fishers in Washington

Shively, K.A., Reese, E.M., Ransom, J.I., Lewis, J.C., Chestnut, T., Werntz, D.O., Levi, T., Prugh, L.R.

Animal Conservation

Genetic Diversity, Demographic Parameters, and Trophic Ecology of the Pampas Cat (Leopardus garleppi) in a Ramsar Wetland of Northwestern Peru

Santiago-Plata, M., Adams, J., Rachlow, J.L., Hurtado, C.M., Garcia-Olaechea, A., Levi, T., Waits, L.P.

Genes 17(3):320

Compositions and methods for control of Pseudogymnoascus destructans

Urbina, J., Dziedzic, E., Levi, T.

U.S. Patent Application 19/227,370

The Ecological Impacts of Large-Carnivore Recovery in North America

Wilmers, C.C., Levi, T., Ruprecht, J., Prugh, L.R., Stahler, D.R.

Annual Review of Ecology, Evolution, and Systematics 56:337–363

Niche partitioning among Neotropical felids

Dymit, E., Twining, J., Garcia-Anleu, R., Levi, T.

Journal of Animal Ecology

From coastlines to mountain crests – regional and continental variation in North American marten diet and foraging

Martin, M., Delheimer, M., Roddy, A.M., Moriarty, K., Eriksson, C., Szykman Gunter, M., Hartman, J., Smith, B., Levi, T.

Ecosphere

Integrating spatial and behavioral data provides comprehensive assessment of grizzly bear-ecotourism coexistence in Nuxalk Territory

Field, K.A., Moody, J.E., Clapham, M., Clark, D.A., Khan, P.B., Levi, T., Paquet, P.C., Darimont, C.T.

Nature Scientific Reports

Snow avalanches and the impact of climate-linked extreme events on mountain ungulate population dynamics and resilience

White, K.S., Levi, T., Hood, E., Darimont, C.T.

Global Change Biology

Life-history trade-offs and environmental variability shape reproductive demography in a mountain sentinel

White, K.S., Levi, T., Hood, E., Darimont, C.T.

Journal of Animal Ecology

Wild canids and felids differ in their reliance on reused travel routeways

Fagan, W.F., et al.

Proceedings of the National Academy of Sciences

Developing custom computer vision models with Njobvu-AI: A collaborative, user-friendly platform for ecological research

Appel, C.L., Subramanian, A., Koning, J.S., Ngosi, M., Sullivan, C.M., Levi, T., Lesmeister, D.B.

Ecological Applications

White-nose syndrome surveillance and bat monitoring activities in North Coast and Cascades network parks 2016–2024

Chestnut, T., Urbina, J., Hansen, M.E., McCaffery, R., Rhea-Fournier, D.J., Allen, J., Levi, T.

Science Report NPR/SR-2025/340, National Park Service

Avoidance-attraction ratios incorrectly characterize behavioral interactions with camera trap data

Dymit, E., Garcia-Anleu, R., Levi, T.

Ecology

Variability and correlations among vital rates and their influence on population growth in mule and black-tailed deer

Ruprecht, J., Forrester, T.D., Clark, D.A., Wisdom, M.J., Smith, J.B., Levi, T.

Journal of Wildlife Management

Wildfire and drought alter the ecology of jaguars and co-occurring mammals in the world's largest wetland

Eriksson, C.E., Kantek, D.L.Z., Miyazaki, S.S., Barry, B.R., Muniz, C.C., dos Santos-Filho, M., Peres, C.A., Levi, T.

Global Change Biology

Using iDNA to determine impacts of Amazonian deforestation on Leishmania hosts, vectors, and their interactions

Massey, A.L., Silva, D.J.F.d., Vieira, C.J.S.P., Allen, J., Canale, G.R., Bernardo, C.S.S., Bronzoni, R.V.d.M., Peres, C.A., Levi, T.

PLOS Neglected Tropical Diseases

Decoding owl calls: Refining occupancy inference from passive acoustic monitoring

Rugg, N.M., Appel, C.L., Jenkins, J.M.A., McCafferty, C., Levi, T., Lesmeister, D.B.

Ecology and Evolution

Landscape heterogeneity shapes the spatial and diet partitioning of a montane carnivore guild

Martin, M., Matthews, S., Stock, S., et al.

Oikos

Bobcat Predation on American Alligators in Coastal South Carolina

Keating, M.P., Rainwater, T.R., Singh, R., Priore, M.R., Platt, S.G., Wilkinson, P.M., Levi, T., Jachowski, C.M.B., Jachowski, D.S.

Southeastern Naturalist

Grizzly bears detected at ecotourism sites are less likely than predicted by chance to encounter conflict

Field, K.A., Moody, J.E., Levi, T., Darimont, C.T.

Canadian Journal of Zoology

Describing diet of imperiled Sierra Nevada red foxes and a carnivoran competitor using DNA metabarcoding

Delheimer, M.S., Martin, M.E., Hartman, J., Moriarty, K.M., Allen, J.M., Levi, T.

Ecology and Evolution

Large-scale overlap and fine-scale avoidance: assessing interactions between coyotes, bobcats, and cougars at multiple scales

Jensen, R., Ruprecht, J., Orning, E., Levi, T., Clark, D.E., Forrester, T.D.

Journal of Mammalogy 105(4):706–717

Molecular and spatial evaluation of small Rodents and Didelphimorphis infected with Mycobacterium leprae in the Southern Amazon, Brazil

Baptista, I., Valois, E., Oliveira, N., et al.

Zoonoses and Public Health

Mule deer selection of fuel reductions is restricted by site fidelity and structured by circadian and seasonal patterns

Forrester, T.D., Spitz, D.B., Merems, J.L., Wisdom, M.J., Rowland, M.M., Johnson, B.K., Smith, J.B., Levi, T., Long, R.A., Clark, D.A.

Forest Ecology and Management

Seek, and you will find: cryptic diversity of the cardiopulmonary nematode Angiostrongylus vasorum in the Americas

Rojas, A., Robleto-Quesada, J., Umaña-Blanco, A.S., Allen, J., Levi, T.

Acta Tropica

Western spotted skunk spatial ecology in the temperate rainforests of the Pacific Northwest

Tosa, M., Lesmeister, D.B., Levi, T.

Ecosphere

Predation risk for hoary marmots in Washington's North Cascades Mountains

Whiles, L., Shipley, L.A., Akins, J.R., Christophersen, R.G., Ransom, J.I., Levi, T., Thornton, D.H.

Journal of Mammalogy

Hunting and trapping in the Americas: the assessment and projection of harvest on wildlife populations

Levi, T., Peres, C.A., Shepard, G.H.

in Johnson, E.A. editor. Natural science and indigenous knowledge. Cambridge University Press

The origin, connectivity, and individual specialization of island wolves after deer extirpation

Eriksson, C.E., Roffler, G.H., Allen, J.M., Lewis, A., Levi, T.

Ecology and Evolution

The effects of both habitat selection and avoidance of competitors on carnivore interactions

Jensen, R., Ruprecht, J.S., Orning, E.K., Levi, T., Clark, D.A., Forrester, T.D.

Journal of Mammalogy

Combining environmental DNA and remote sensing for efficient fine-scale mapping of arthropod biodiversity

Yuanheng, L., Devenish, C., Tosa, M., Luo, M., Bell, D., Lesmeister, D.B., Greenfield, P., Pichler, M., Levi, T., Yu, D.W.

Philosophical Transactions of the Royal Society B

Human disturbance in riparian areas disrupts predator-prey interactions between grizzly bears and salmon

Adams, M.S., Levi, T., Bourbonnais, M., Service, C., Artelle, K., Bryan, H., Paquet, P., Nelson, T., Darimont, C.

Ecology and Evolution

Intraguild predation of cougar kittens following the aggregation of American black bears

Bianco, M.D., Ruprecht, J., Clark, D.A., Forrester, T.D., Levi, T.

Ecosphere

Recovery of marine keystone predator transforms terrestrial predator–prey dynamics

Roffler, G.H., Eriksson, C.E., Allen, J.M., Levi, T.

Proceedings of the National Academy of Sciences

Warm oceans exacerbate Chinook salmon bycatch in the Pacific hake fishery driven by thermal and diel depth-use behaviors

Sabal, M., Richerson, K., Moran, P., Levi, T., Tuttle, V.J., Banks, M.

Fish and Fisheries 24(6):910–923

Humanity's diverse predatory niche and its ecological consequences

Darimont, C.T., Cooke, R., Bourbonnais, M.L., Bryan, H.M., Carlson, S.M., Estes, J.A., Galetti, M., Levi, T., MacLean, J.L., McKechnie, I., Paquet, P.C., Worm, B.

Communications Biology 6:1211

Thermal refuge use and parasitism: spatiotemporal variation in anchor worm and lamprey wounds on Klamath redband trout

Ortega, J., Hahlbeck, N., Derrickson, C., et al.

Ecosphere

Teasing apart the dilution effect by combining DNA metabarcoding and statistical modeling

Levi, T., Massey, A.L.

Molecular Ecology

Using passive acoustic monitoring to estimate Northern Spotted Owl landscape use and pair occupancy

Appel, C.L., Lesmeister, D.B., Duarte, A., Davis, R.J., Weldy, M.J., Levi, T.

Ecosphere 14(2):e4421

Density-dependent changes in elk resource selection following timber harvest over successional time scales

Ruprecht, J., Wisdom, M., Clark, D., Rowland, M., Levi, T.

Ecological Applications

Variation in adult and pup wolf diets at natal den sites is influenced by forest composition and configuration

Roffler, G.H., Levi, T., Pilgrim, K., Zarn, K.E., Schwartz, M.K.

Ecology and Evolution

Creating, curating, and evaluating a mitogenomic reference database to improve regional species identification using environmental DNA

Dziedzic, E., Sidlauskas, B., Cronn, R., Anthony, J., Cornwell, T., Friesen, T., Konstantinidis, P., Penaluna, B., Stein, S., Levi, T.

Molecular Ecology Resources

A seasonal pulse of ungulate neonates influences predator space use in a multipredator-multiprey system

Ruprecht, J., Forrester, T.D., Jackson, N.J., et al.

Ecology and Evolution

Multi-locus DNA metabarcoding reveals seasonality of foraging ecology of western spotted skunks in the Pacific Northwest

Tosa, M., Lesmeister, D.B., Levi, T.

Ecosphere

Njobvu-AI: An open-source tool for collaborative image labeling and implementation of computer vision models

Koning, J.S., Subramanian, A., Alotaibi, M., et al.

ArXiv

Barred owl predation of western spotted skunks

Tosa, M., Lesmeister, D., Levi, T.

Northwestern Naturalist

The roles of humans and apex predators in sustaining ecosystem structure and function

Lennox, R.J., Brownscomb, J.W., Darimont, C.T., Horodysky, A., Levi, T., Raby, G.D., Cooke, S.J.

Ecology and Society

The spatial overlap of small-scale cannabis farms with aquatic and terrestrial biodiversity

Parker-Shames, P., Choi, C., Bustic, V., Green, D., Barry, B., Moriarty, K.M., Levi, T., Brashares, J.

Conservation Science and Practice

Invertebrates for vertebrate biodiversity monitoring: comparisons using three insect taxa as iDNA samplers

Massey, A.L., Bronzoni, R.V.d.M., Silva, D.J.F.d., Allen, J., Lazari, P.R.d., Filho, M.d.S., Canale, G.R., Peres, C.A., Levi, T.

Molecular Ecology Resources

The multivariate Tweedie: A practical primer on generalized linear models for diet analysis derived from foraging theory

Thorson, J.T., Arimitsu, M.L., Levi, T., Roffler, G.H.

Ecology

Extensive aquatic subsidies lead to territorial breakdown and high density of an apex predator

Eriksson, C.E., Kantek, D.L., Miyazaki, S.S., Morato, R.G., dos Santos-Filho, M., Ruprecht, J.S., Peres, C.A., Levi, T.

Ecology

Land use effects on mosquito biodiversity and potential arbovirus emergence in the Southern Amazon, Brazil

Vieira, C.J.S.P., Bernardo, C.S.S., Silva, D.J.F., et al.

Transboundary and Emerging Diseases

Integrating multi-method surveys and recovery trajectories into species distribution models

Barry, B.R., Moriarty, K.M., Green, D., Hutchinson, R., Levi, T.

Ecosphere

The Rapid Rise of Next-Generation Natural History

Tosa, M.I., Dziedzic, E.H., Appel, C.L., Urbina, J., Massey, A., Ruprecht, J., Eriksson, C.E., Dolliver, J.E., Lesmeister, D.B., Betts, M.G., Peres, C.A., Levi, T.

Frontiers in Ecology and Evolution p.480

Wolf dietary diversity in an island archipelago

Roffler, G.H., Allen, J.M., Massey, A.L., Levi, T.

Bulletin of the Ecological Society of America 102(1):1–6

Predicted distributions of Humboldt martens: historical and current context

Moriarty, K.M., Linnell, M., Thompson, J., et al.

PeerJ

Variable strategies to solve risk-reward tradeoffs in carnivore communities

Ruprecht, J.S., Eriksson, C.E., Forrester, T.D., Spitz, D., Clark, D.A., Wisdom, M., Rowland, M., Bianco, M., Smith, J., Levi, T.

Proceedings of the National Academy of Sciences

Integrating spatial capture-recapture models with variable individual identification

Ruprecht, J.S., Eriksson, C.E., Forrester, T.D., Clark, D.A., Wisdom, M.J., Rowland, M.M., Johnson, B.K., Levi, T.

Ecological Applications

Building intuition regarding the statistical behavior of mass medical testing programs

Waller, L., Levi, T.

Harvard Data Science Review

Local values and data empower culturally-guided and ecosystem-based fisheries management

Adams, M.S., Connors, B., Levi, T., Shaw, D., Walkus, J., Rogers, S., Darimont, C.T.

Marine and Coastal Fisheries

Comparison of mechanical sorting and DNA metabarcoding for diet analysis with degraded wolf scats

Massey, A.L., Roffler, G., Vermeul, T., Allen, J.M., Levi, T.

Ecosphere

Fisher recolonization and recovery in Oregon: Feasibility assessment for the reintroduction of fishers

Matthews, S.M., Green, D.S., Martin, M.E., et al.

Mesocarnivore landscape use along a gradient of urban, rural, and forest cover

Rodriguez, J.T., Lesmeister, D.B., Levi, T.

PeerJ

Pseudogymnoascus destructans growth in wood, soil and guano substrates

Urbina, J., Chestnut, T., Allen, J., Levi, T.

Nature Scientific Reports

A forest loss report card for the world's protected areas

Wolf, C., Levi, T., Ripple, W.J., Zárrate-Charry, D.A., Betts, M.G.

Nature Ecology and Evolution

Diet of invasive wild pigs in a landscape dominated by sugar cane plantations

Pedrosa, F., Berce, W., Costa, V.E., Levi, T., Galetti, M.

Journal of Mammalogy

Regional metabarcoding of fecal DNA shows that dietary diversification in wolves substitutes for ungulates in an island archipelago

Roffler, G.H., Allen, J.M., Massey, A.L., Levi, T.

Ecosphere

Interaction ecology and conservation of bear-salmon ecosystems

Levi, T., Hilderbrand, G.V., Hocking, M.D., et al.

Frontiers in Ecology and Evolution

Integrating genetic data and demographic modeling to facilitate conservation of small isolated mountain goat populations

White, K.W., Levi, T., Breen, J., et al.

Journal of Wildlife Management

Better Boundaries: Environmental DNA improves detection of the upper limit of fish distribution in forested streams

Penaluna, B.E., Allen, J.M., Arismendi, I., Levi, T., Garcia, T.S., Walter, J.

Ecosphere

More affordable and effective noninvasive SNP genotyping using high-throughput amplicon sequencing

Eriksson, C.E., Ruprecht, J., Levi, T.

Molecular Ecology Resources

Individual variation in use of the nutritional landscape predicts early-winter condition of a long-lived ungulate

Merems, J.L., Shipley, L.A., Levi, T., Ruprecht, J., Clark, D.A., Wisdom, M.J., Jackson, N.J., Stewart, K.M., Long, R.A.

Frontiers in Ecology and Evolution

Environmental DNA facilitates accurate, inexpensive, and multi-year population estimates of millions of anadromous fish

Pochardt, M., Allen, J.M., Hart, T., Miller, S.D., Yu, D.W., Levi, T.

Molecular Ecology Resources

Orthohantavirus antibodies among phyllostomid bats from the arc of deforestation in Southern Amazonia, Brazil

Sabino-Santos, G., Fernandes, F.F., et al.

Transboundary and Emerging Diseases

Experimental evaluation of degradation rates of genomic DNA of the pathogen Pseudogymnoascus destructans in bat guano

Urbina, J., Chestnut, T., Schwalm, D., Allen, J., Levi, T.

PeerJ

Intrapopulation isotopic niche diversity over landscapes: spatial patterns inform conservation of bear-salmon systems

Adams, M.S., Service, C., Bateman, A., et al.

Ecosphere

Intraspecific variation in movement strategies among bald eagles (Haliaeetus leucocephalus) in an anadromous fish system

Wheat, R.E., Lewis, S.B., Wang, Y., Levi, T., Wilmers, C.C.

Movement Ecology

Extinction filters mediate the global effects of fragmentation

Betts, M.G., Wolf, C., et al.

Science

Eating plants and planting trees for climate mitigation

Wolf, C., Ripple, W.J., Betts, M.G., Levi, T., Peres, C.A.

Global Change Biology

Biotic factors influencing the unexpected distribution of a Humboldt marten population in forested sand dunes

Eriksson, C.E., Moriarty, K.M., Linnell, M.A., Levi, T.

PLoS ONE

Non-consumptive behavioral responses and nutritional consequences of elk when avoiding human hunters

Spitz, D.B., Rowland, M., Clark, D.A., Wisdom, M.J., Smith, J.B., Brown, C.L., Levi, T.

Ecosphere

Seed dispersal effectiveness by a large-bodied invasive species in defaunated landscapes

Pedrosa, F., Bercê, W., Levi, T., Pires, M., Galetti, M.

Biotropica

Tropical forests can maintain hyperdiversity because of enemies

Levi, T., Barfield, M., Barrantes, S., Sullivan, C., Holt, R.D., Terborgh, J.

Proceedings of the National Academy of Sciences

Reply to Cannon and Lerdau: Maintenance of tropical forest tree diversity

Levi, T., Barfield, M., Holt, R.D., Terborgh, J.

Proceedings of the National Academy of Sciences

Environmental DNA for the enumeration and management of Pacific salmon

Levi, T., Allen, J.M., Bell, D., Joyce, J., Russell, J.R., Tallmon, D.A., Vulstek, S.C., Yahan, Y., Yu, D.W.

Molecular Ecology Resources

Tick-borne disease risk in a forest food web

Ostfeld, R.S., Levi, T., Keesing, F., Oggenfuss, K., Canham, C.D.

Ecology

Fire history influences large-herbivore behavior at circadian, seasonal, and successional scales

Spitz, D., Clark, D., Wisdom, M.J., Rowland, M., Johnson, B., Long, R.A., Levi, T.

Ecological Applications

Large species within Carnivora are large carnivores

Wolf, C., Betts, M.G., Levi, T., Newsome, T.M., Ripple, W.J.

Royal Society Open Science

DNA metabarcoding reveals human-induced shifts in dietary niche partitioning in a carnivore guild

Smith, J.A., Thomas, A.C., Levi, T., Wang, Y., Wilmers, C.C.

Oikos

Density and population trajectories of coastal marten: a rare and geographically isolated small carnivore

Linnell, M.A., Moriarty, K.M., Green, D.S., Levi, T.

PeerJ

Seed dispersal and resource subsidies from salmon-supported bears to granivores

Shakeri, Y., White, K.W., Levi, T.

Ecosphere

The primacy of bears as seed dispersers in salmon-bearing ecosystems

Harrer, L.E.F., Levi, T.

Ecosphere

Projecting the future of an alpine ungulate under climate change scenarios

White, K.S., Gregovic, D., Levi, T.

Global Change Biology

Does biodiversity protect humans against infectious disease?: Comment

Levi, T., Massey, A.L., Holt, R.D., Keesing, F., Ostfeld, R.S., Peres, C.A.

Ecology 97:536–546

Global forest loss disproportionately erodes biodiversity in intact landscapes

Betts, M.G., Wolf, C., Ripple, W.J., et al.

Nature

Lyme disease ecology in a changing world: consensus, uncertainty, and critical gaps for improving control

Kilpatrick, A.M., Dobson, A.D.M., Levi, T., Salkeld, D.J., Swei, A., Ginsberg, H.S., Kjemtrup, A., Padgett, K.A., Jensen, P.M., Fish, D., Ogden, N.H., Diuk-Wasser, M.

Philosophical Transactions of the Royal Society B

Conserving the World's Megafauna and Biodiversity: The Fierce Urgency of Now

Ripple, W.J., Chapron, G., Lopez-Bao, J.V., et al.

BioScience

The impact of temperature and precipitation during Ixodes scapularis questing periods on human Lyme disease incidence

Burtis, J.C., Sullivan, P., Ostfeld, R.S., Levi, T., Yavitt, J.B., Fahey, T.J.

Parasites and Vectors

Environmental DNA from residual saliva for efficient noninvasive genetic monitoring of brown bears (Ursus arctos)

Wheat, R.E., Allen, J.M., Miller, S.D.L., Wilmers, C.C., Levi, T.

PLoS ONE

Are we eating the world's mammals to extinction?

Ripple, W.J., Abernethy, K., Chapron, G., Levi, T., et al.

Royal Society Open Science

Empty rivers rather than empty forests: a century of commercial hunting in Amazonia

Antunes, A.P., Fewster, R., Venticinque, E.M., Peres, C.A., Levi, T., Rohe, F., Shepard, G.H.

Science Advances

Line transect surveys under detect terrestrial mammals: implications for the sustainability of subsistence hunting

Fragoso, J.M.V., Levi, T., Oliveira, L.F.B., et al.

PLoS ONE

Dispersal limitation induces long-term biomass collapse in overhunted Amazonian forests

Peres, C.A., Emilio, T., Schietti, J., Desmouliere, S.J.M., Levi, T.

Proceedings of the National Academy of Sciences 113:892–897

Long-term aspen dynamics and trophic cascades in Northern Yellowstone

Beschta, R.L., Painter, L.E., Levi, T., Ripple, W.J.

Canadian Journal of Forest Research 46:548–556

Quantifying dilution and amplification in a community of hosts for tick-borne pathogens

Levi, T., Keesing, F., Holt, R.D., Barfield, M., Ostfeld, R.S.

Ecological Applications 26(2):285–294

Differential use of salmon by vertebrate consumers: implications for conservation

Levi, T., Wheat, R., Allen, J.M., Wilmers, C.C.

PeerJ 3:e1157

Collapse of the World's Largest Herbivores

Ripple, W.J., Newsome, T.M., Wolf, C., et al.

Science Advances 1(4):e1400103

Threshold levels of generalist predation determine consumer response to resource pulses

Levi, T., Kilpatrick, A.M., Barfield, M., Holt, R.D., Mangel, M., Wilmers, C.C.

Oikos

Accelerated phenology of blacklegged ticks under climate warming

Levi, T., Keesing, F., Oggenfuss, K., Ostfeld, R.S.

Philosophical Transactions of the Royal Society B 370:20130556

Saving the world's terrestrial megafauna

Ripple, W.J., et al.

BioScience 65(10):1117–1124

Working together: A call for inclusive conservation

Tallis, H., et al.

Nature 515:27–28

Life history and demographic drivers of reservoir competence for three tick-borne zoonotic pathogens

Ostfeld, R.S., Levi, T., Jolles, A., Martin, L.B., Hosseini, P.R., Keesing, F.

PLoS ONE 9(9):e107387

Several scales of biodiversity affect ecosystem multifunctionality

Pasari, J., Levi, T., Zavaleta, E., Tilman, D.

Proceedings of the National Academy of Sciences 110:10219–10222

Reply to Mokany et al: Comprehensive measures of biodiversity are critical to investigations of ecosystem multifunctionality

Pasari, J., Levi, T., Zavaleta, E., Tilman, D.

Proceedings of the National Academy of Sciences 110(43)

Competition and facilitation in the capuchin-squirrel monkey relationship

Levi, T., Silvius, K.M., Oliveira, L.F.B., Fragoso, J.M.V.

Biotropica

Dispersal vacuum in the seedling recruitment of a primate-dispersed Amazonian tree

Levi, T., Peres, C.A.

Biological Conservation 163:99–106

Do irrigation and predator control reduce the productivity of migratory ungulate herds?

Wilmers, C.C., Levi, T.

Ecology 94:1264–1270

Climate and vegetation phenology: Predicting the effects of warming temperatures

Wilmers, C.C., Ram, K., Watson, F.G.R., White, P.J., Smith, D.W., Levi, T.

in White, P.J. editor. Yellowstone's Wildlife in Transition. Harvard University Press

Reliable, verifiable, and efficient monitoring of biodiversity via metabarcoding

Ji, Y., Ashton, L., Pedley, S.M., et al.

Ecology Letters 16(10):1245–1257

Resolviendo el conflicto "parque-personas" en el Manu

Yu, D.W., Shepard, G.H., Ohl-Schacherer, J., Levi, T.

in Groenendijk, J. and A. Tovar editors. Reporte Manu. San Diego Zoo Global Peru y SERNANP

Deer, predators, and the emergence of Lyme disease

Levi, T., Kilpatrick, A.M., Mangel, M., Wilmers, C.C.

Proceedings of the National Academy of Sciences 109:10942–10947

Using Grizzly bears to assess harvest-ecosystem tradeoffs in salmon fisheries

Levi, T., Darimont, C.T., MacDuffee, M., Mangel, M., Paquet, P., Wilmers, C.C.

PLoS Biology 10(4)

Wolves-Coyotes-Foxes: A cascade among carnivores

Levi, T., Wilmers, C.C.

Ecology 93:921–929

Hunting in ancient and modern Amazonia: Rethinking sustainability

Shepard, G.H., Levi, T., Neves, E., Peres, C.A., Yu, D.W.

American Anthropologist 114(4)

The behavior and diet breadth of central-place foragers: an application to human hunters and Neotropical game management

Levi, T., Lu, F., Yu, D.W., Mangel, M.

Evolutionary Ecology Research 13:171–185 (2011)

Spatial tools for modeling the sustainability of subsistence hunting in tropical forests

Levi, T., Shepard, G.H., Ohl-Schacherer, J., Wilmers, C.C., Peres, C.A., Yu, D.W.

Ecological Applications 21:1802–1818 (2011)

Conservation in Low-Governance Environments

Yu, D.W., Levi, T., Shepard, G.H.

Biotropica 42:569–571 (2010)

Modeling the long-term sustainability of indigenous hunting in Manu National Park, Peru

Levi, T., Shepard, G.H., Ohl-Schacherer, J., Peres, C.A., Yu, D.W.

Journal of Applied Ecology 46:804–814 (2009)

The sustainability of hunting by Matsigenka native communities in Manu National Park, Peru

Ohl-Schacherer, J., Shepard, G.H., Kaplan, H., Peres, C.A., Levi, T., Yu, D.W.

Conservation Biology 21:1174–1185 (2007)